Replace the (?)

The below given figure comprises of a pattern through which you can determine the missing letter. Can you push your mind to find the pattern and add the missing letter?

Replace the (?)




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In 2011, people playing Foldit, an online puzzle game about protein folding, resolved the structure of an enzyme that causes an Aids-like disease in monkeys. Researchers had been working on the problem for 13 years. The gamers solved it in three weeks.